“Dryophyllum” leaf forms and stomata


A study of the “Dryophyllum” leaf forms from the Palaeocene of Southeastern North America

by Jones J. H., Dilcher D. L. (1988)

Jay H. Jones, Biology Department, University of La Verne, USA

David L. Dilcher, Indiana University, Bloomington, Indiana, U.S.A

in Palaeontographica Abt. B, 208 (4-8),: 53-80


“Dryophyllum” leaf forms are ‘among the most abundant in the Paleogene of southeastern North America. These leaf types along with other related leaf forms have been thoroughly reexamined using modern methods of foliar analysis. The “Dryophyllum” leaf forms fall into two groups both of which cannot be accommodated in the genus Dryophyllum DEBEY ex SAPORTA.

The genus Castaneophyllum is erected to hold the castaneoid leaf forms previously assigned to Dryophyllum tennesseense and D. moorii. The second group, which contains leaf forms previously assigned to D. puryearense, Banksia saffordii and Banksia tenuifolia has been assigned to the new genus Berryophyllum.

Members of the genus Castaneophyllum seem to be well within the bounds of the Castaneoideae and may represent species of the modern genus Castanea.

Members of the genus Berryophyllum although closest to the Quercoideae are not very similar to any modern Fagaceae. The combination of foliar characters suggest that Berryophyllum may be ancestral within the family.

Description of the stomata in the analysis of the different groups.


Plate 2, Fig. 13: Photomicrograp of lower Castaneophyllum tennesseense cuticle showing anomocytic stomatal complex and septate trichome base (NL 2986) 650x.

Communicated by Parisa Panahi

Published by

Willem Van Cotthem

Honorary Professor of Botany, University of Ghent (Belgium). Scientific Consultant for Desertification and Sustainable Development.

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