Stomata in Litsaea

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Litsea cubeba

Study of Stomata in Fifteen Species of Litsaea Lamk. of Family Lauraceae

by Vaidya M. (2016)

Meenakshi VaidyaMeenakshi_Vaidya2

in Asian Journal of Biochemical and Pharmaceutical Research Issue 1 (Vol. 6) 2016 ISSN: 2231-2560 –

http://ajbpr.com/issues/volume6/issue1/FINAL%201.pdf 

Abstract:

Cuticular features of Lauraceae were studied in the early twentieth century. Litsaea Lamk. belongs to family Lauraceae.

The mature stomata of 15 species of Litsaea are studied. The leaves are hypostomatic. The stomata are absent on the upper epidermis. The stomata are present only on the lower epidermis.

The types of stomata observed in the studied species of Litsaea are paracytic, anomocytic, anisocytic, brachyparacytic, hemiparacytic, brachyparatetracytic and amphiparacytic type.

These features help in the identification of the species as the morphological key of this genus is based on development of anther. Thus we can make use of anatomy as a tool for the identification of the species of this genus Litsaea.

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OBSERVATIONS:

Litsaea Meissneri: Upper epidermis (Fig. 1): Stomata are absent on the upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 2): Stomata are anisocytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular. The epidermal cells are polygonal in shape.

Litsaea nitida: Upper epidermis (Fig. 3): Stomata are absent on the upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 4): Stomata are paracytic type. The guard cells are elongated and kidney shaped, epidermal cells are irregular. The epidermal cells are polygonal.

Litsaea oblonga: Upper epidermis (Fig. 5): Stomata are absent on the upper epidermis. The epidermal cells are irregular. The outline of the epidermal cells is slightly wavy.

Lower epidermis (Fig. 6): Stomata are anomocytic type. The guard cells are elongatd and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea oleoides: Upper epidermis (Fig. 7): Stomata are absent on the upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 8): Stomata are anomocytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular.

Litsaea Panamonja: Upper epidermis (Fig. 9): Stomata are absent on upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 10): Stomata are anomocytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular and polygonal in shape.

Litsaea polyantha: Upper epidermis (Fig. 11): Stomata are absent on upper epidermis. The epidremal cells are irregular and polygonal.

Lower epidermis (Fig. 12): Stomata are anomocytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular and polygonal in shape.

Litsaea salicifolia: Upper epidermis (Fig. 13): Stomata are absent on the upper epidermis. Epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 14): Stomata are anomocytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular and polygonal in shape.

Litsaea scrobiculata: Upper epidermis (Fig. 15): Stomata are absent on the upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 16): Stomata are anomocytic type. The guard cells are elongated and kidney shaped. Subsidiary cells are not distinguishable from epidermal cells. The epidermal cells are irregular and polygonal in shape.

2

Asian Journal of Biochemical and Pharmaceutical Research Issue 1 (Vol. 6) 2015 ISSN: 2231-2560 CODEN (USA): AJBPAD

Litsaea sebifera: Upper epidermis (Fig. 17): Stomata are absent on upper epidermis. The epidermal cells are irregular. The epidermal cells have a wavy wall.

Lower epidermis (Fig. 18): Stomata are paracytic and brachyparacytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea semecarpifolia: Upper epidermis (Fig. 19): Stomata are absent on upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 20): Stomata are paracytic type. The guard cells are elongated and kidney shaped. Epidermal cells are irregular and polygonal in shape.

Litsaea stocksii: Upper epidermis (Fig. 21): Stomata are absent on upper epidermis. The epidemal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 22): Stomata are paracytic and brachyparatetracytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea Thomsonii: Upper epidermis (Fig. 23): Stomata are absent on the upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 24): The stomata are paracytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea umbrosa: Upper epidermis (Fig. 25): Stomata are absent on upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 26): The stomata are paracytic and amphiparacytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea Wightiana: Upper epidermis (Fig. 27): The stomata are absent on upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 28): The stomata are paracytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Litsaea zeylanica: Upper epidermis (Fig. 29): The stomata are absent on upper epidermis. The epidermal cells are irregular and polygonal in shape.

Lower epidermis (Fig. 30): The stomata are paracytic and hemiparacytic type. The guard cells are elongated and kidney shaped. The epidermal cells are irregular and polygonal in shape.

Anomocytic type of stomata are observed in Litsaea oblonga, L. oleoides, L. Panamonja, L. polyantha, L. salicifolia and L. scrobiculata.

Paracytic type of stomata are observed in Litsaea nitida, L. semecarpifolia, L. Thomsonii and L. Wightiana.

Anisocytic type of stomata are observed in Litsaea Meissneri.
Paracytic and Brachyparacytic type of stomata are observed in Litsaea sebifera. Paracytic and Brachyparatetracytic type of stomata are observed in Litsaea stocksii. Paracytic and Hemiparacytic type of stomata are observed in Litsaea zeylanica.

Paracytic and Amphiparacytic type of stomata are observed in Litsaea umbrosa. 3

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Published by

Willem Van Cotthem

Honorary Professor of Botany, University of Ghent (Belgium). Scientific Consultant for Desertification and Sustainable Development.

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